@article {Burkett2016,
	title = {{Colonization of over a thousand Cibicidoides wuellerstorfi (foraminifera: Schwager, 1866) on artificial substrates in seep and adjacent off-seep locations in dysoxic, deep-sea environments}},
	journal = {Deep Sea Research Part I: Oceanographic Research Papers},
	volume = {117},
	year = {2016},
	pages = {39{\textendash}50},
	abstract = {After {\~{}}1 yr on the seafloor at water depths of {\~{}}700 m on Hydrate Ridge in the Pacific, eight colonization experiments composed primarily of a plastic mesh cube (from here on refered to as SEA3, for Seafloor Epibenthic Attachment Cubes) were colonized by 1076 Cibicidoides wuellerstorfi on {\~{}}1841 cm2 of experimental substrate. This species is typically considered an indicator of well-oxygenated conditions, and recruitment of such large numbers in bottom waters with low dissolved oxygen availability (0.24{\textendash}0.37 mL/L) indicate that this taxon may not be as limited by oxygen as previously thought. Clues about substrate preferences were evident from the distribution, or lack thereof, of individuals among plastic mesh, coated steel frame, wooden dowels and reflective tape. Abundance, individual size distributions within cage populations and isotopic biogeochemistry of living foraminifera colonizing experimental substrates were compared between active seep and adjacent off-seep experiment locations, revealing potential differences between these environments. Few studies have examined foraminiferal colonization of hard substrates in the deep-sea and to our knowledge no previous study has compared foraminiferal colonization of active seep and off-seep substrates from the same region. This study provides initial results of recruitment, colonization, geochemical and morphological aspects of the paleoceanographically significant species, C. wuellerstorfi, from dynamic deep-sea environments. Further experimental deployments of SEA3s will provide a means to assess relatively unknown ecologic dynamics of important foraminiferal deep-sea species.},
	keywords = {HOV Alvin (Human Occupied Vehicle), ROV Jason (Remotely Operated Vehicle)},
	doi = {10.1016/j.dsr.2016.08.011},
	author = {Burkett, A M and Rathburn, A E and Perez, Elena M and Levin, L A and Martin, J B}
}
@article {Guilini2012,
	title = {{Cold seep and oxygen minimum zone associated sources of margin heterogeneity affect benthic assemblages, diversity and nutrition at the Cascadian margin (NE Pacific Ocean)}},
	journal = {Progress in Oceanography},
	volume = {96},
	year = {2012},
	pages = {77{\textendash}92},
	abstract = {Hydrate Ridge (HR), located on the northeastern Pacific margin off Oregon, is characterized by the presence of outcropping hydrates and active methane seepage. Additionally, permanent low oxygen conditions overlay the benthic realm. This study evaluated the relative influence of both seepage and oxygen minima as sources of habitat heterogeneity and potential stress-inducing features on the bathyal metazoan benthos (primarily nematodes) at three different seep and non-seep HR locations, exposed to decreasing bottom-water oxygen concentrations with increasing water depth. The nematode seep communities at HR exhibited low diversity with dominance of only one or two genera (Daptonema and Metadesmolaimus), elevated average individual biomass and $δ$13C evidence for strong dependance on chemosynthesis-derived carbon, resembling deep-sea seeps worldwide. Although the HR seep habitats harbored a distinct nematode community like in other known seep communities, they differed from deep-sea seeps in well-oxygenated waters based on that they shared the dominant genera with the surrounding non-seep sediments overlain by oxygen-deficient bottom water. The homogenizing effect of the oxygen minimum zone on the seep nematode assemblages and surrounding sediments was constant with increasing water depth and concomitant greater oxygen-deficiency, resulting in a loss of habitat heterogeneity.},
	keywords = {HOV Alvin (Human Occupied Vehicle)},
	issn = {0079-6611},
	doi = {10.1016/j.pocean.2011.10.003},
	url = {http://dx.doi.org/10.1016/j.pocean.2011.10.003},
	author = {Guilini, K and Levin, L A and Vanreusel, A}
}
@article {Bernardino2012,
	title = {{Comparative Composition, Diversity and Trophic Ecology of Sediment Macrofauna at Vents, Seeps and Organic Falls}},
	journal = {PLoS ONE},
	volume = {7},
	number = {4},
	year = {2012},
	month = {apr},
	pages = {e33515},
	publisher = {Public Library of Science},
	abstract = {Sediments associated with hydrothermal venting, methane seepage and large organic falls such as whale, wood and plant detritus create deep-sea networks of soft-sediment habitats fueled, at least in part, by the oxidation of reduced chemicals. Biological studies at deep-sea vents, seeps and organic falls have looked at macrofaunal taxa, but there has yet to be a systematic comparison of the community-level attributes of sediment macrobenthos in various reducing ecosystems. Here we review key similarities and differences in the sediment-dwelling assemblages of each system with the goals of (1) generating a predictive framework for the exploration and study of newly identified reducing habitats, and (2) identifying taxa and communities that overlap across ecosystems. We show that deep-sea seep, vent and organic-fall sediments are highly heterogeneous. They sustain different geochemical and microbial processes that are reflected in a complex mosaic of habitats inhabited by a mixture of specialist (heterotrophic and symbiont-associated) and background fauna. Community-level comparisons reveal that vent, seep and organic-fall macrofauna are very distinct in terms of composition at the family level, although they share many dominant taxa among these highly sulphidic habitats. Stress gradients are good predictors of macrofaunal diversity at some sites, but habitat heterogeneity and facilitation often modify community structure. The biogeochemical differences across ecosystems and within habitats result in wide differences in organic utilization (i.e., food sources) and in the prevalence of chemosynthesis-derived nutrition. In the Pacific, vents, seeps and organic-falls exhibit distinct macrofaunal assemblages at broad-scales contributing to {\~A}{\"Y} diversity. This has important implications for the conservation of reducing ecosystems, which face growing threats from human activities.},
	doi = {10.1371/journal.pone.0033515},
	url = {http://dx.doi.org/10.1371/journal.pone.0033515},
	author = {Bernardino, A F and Levin, L A and Thurber, A R and Smith, C R}
}
@article {Basak2009,
	title = {{Carbon and oxygen isotope geochemistry of live (stained) benthic foraminifera from the Aleutian Margin and the Southern Australian Margin}},
	journal = {Marine Micropaleontology},
	volume = {70},
	number = {3-4},
	year = {2009},
	pages = {89{\textendash}101},
	keywords = {ROV Jason (Remotely Operated Vehicle)},
	issn = {0377-8398},
	doi = {10.1016/j.marmicro.2008.11.002},
	url = {http://dx.doi.org/10.1016/j.marmicro.2008.11.002},
	author = {Basak, C and Rathburn, A E and Perez, M E and Martin, J B and Kluesner, J W and Levin, L A and De Deckker, P and Gieskes, J M and Abriani, M}
}
@article {Levin2007,
	title = {{Community structure and nutrition of deep methane-seep macrobenthos from the North Pacific (Aleutian) Margin and the Gulf of Mexico (Florida Escarpment)}},
	journal = {Marine Ecology},
	volume = {28},
	number = {1},
	year = {2007},
	pages = {131{\textendash}151},
	keywords = {HOV Alvin (Human Occupied Vehicle)},
	issn = {0173-9565},
	doi = {10.1111/j.1439-0485.2006.00131.x},
	url = {http://dx.doi.org/10.1111/j.1439-0485.2006.00131.x},
	author = {Levin, L A and Mendoza, G F}
}
@article {Levin1991,
	title = {{Control of deep-sea benthic community structure by oxygen and organic-matter gradients in the easter Pacific Ocean}},
	journal = {Journal of Marine Research},
	volume = {49},
	number = {4},
	year = {1991},
	pages = {763{\textendash}800},
	keywords = {HOV Alvin (Human Occupied Vehicle)},
	issn = {0022-2402},
	doi = {10.1357/002224091784995756},
	url = {http://dx.doi.org/10.1357/002224091784995756},
	author = {Levin, L A and Huggett, C L and Wishner, K F}
}
